Publications on Pacific Coleoptera

You can access all my papers on Pacific Beetles via the hyperlinks. Some papers are open access, whereas others may be accessible only if your institution has a subscription. If you hit a publisher paywall and want a copy of the paper, please email me at and I will send you a gratis PDF.

Liebherr, J.K. 2018. Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel. Deutsche Entomologissche Zeitschrift 65(1): 1–63 (

Liebherr, J.K. 2016. Cyphocoleus Chaudoir (Coleoptera: Carabidae: Odacanthini): taxonomic revision, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211–270 (

Liebherr, J.K. 2015. The Mecyclothorax beetles (Coleoptera: Carabidae: Moriomorphini) of Haleakalā, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 522 (6074): 1-407 (DOI: 10.3897/zookeys.544.6074).

Liebherr, J.K. & N. Porch. 2015. Reassembling a lost lowland carabid beetle assemblage (Coleoptera) from Kauai, Hawaiian Islands. Invertebrate Systematics 29: 191-213 (DOI:

Liebherr, J.K. & D.R. Maddison. 2013. Colonization of the Pacific by Bembidion beetles (Coleoptera: Carabidae), with description of Bembidion tahitiense, sp. nov., from Tahiti, French Polynesia. Invertebrate Systematics 27: 439-449 (DOI: 

Liebherr, J.K. 2013. The Mecyclothorax beetles (Coleoptera: Carabidae: Moriomorphini) of Tahiti, Society Islands. ZooKeys 322: 170 pp(DOI:

Liebherr, J.K. 2012. New Mecyclothorax spp. (Coleoptera: Carabidae: Moriomorphini) define Mont Mauru, eastern Tahiti Nui, as a distinct area of endemism. ZooKeys 227: 63-99 (DOI: 10.3897/zookeys.227.3797).

Liebherr, J.K. 2012. The first precinctive Carabidae from Moorea, Society Islands: new Mecyclothorax spp. (Coleoptera) from the summit of Mont Tohiea. ZooKeys 224: 37-80 (DOI:10.3897/zookeys.224.3675).

Liebherr, J.K. & P.D. Krushelnycky. 2011. Mecyclothorax palikea, sp. n. from the Waianae Range, Oahu, and the biogeographical history of Hawaii’s M. flavomarginatus species group (Coleoptera: Carabidae: Moriomorphini). Insect Systematics & Evolution 42(4): 365-384 (×608718).

Liebherr, J.K. 2011. The Mecyclothorax beetles (Coleoptera: Carabidae: Moriomorphini) of West Maui, Hawaii: taxonomy, biogeography, and conservation. Deutsche Entomologische Zeitschrift 58: 15-76 (

Liebherr, J.K., A. Casale, T.L. Erwin & G.E. Ball. 2010. On the provenance of Boheman’s “Eugenies resa” Carabidae (Coleoptera) allegedly described from Hawaii. Coleopterists Bulletin 64: 221-229 (

Liebherr, J.K., S.L. Montgomery, R.A. Englund & G.A. Samuelson. 2009. The first Hawaiian occurrence of the alien ground beetle Agonum muelleri (Coleoptera: Carabidae) from the summit of Mauna Kea, Hawaii island. Proceedings of the Hawaiian Entomological Society 41: 97-103 (

Liebherr, J.K. 2009. Native and alien Carabidae (Coleoptera) share Lanai, an ecologically devastated island. Coleopterists Bulletin 63: 383-411 (

Liebherr, J.K. 2009. Taxonomic revision of the Mecyclothorax beetles (Coleoptera: Carabidae) of Oahu: epithets as epitaphs for an endangered fauna? Systematic Entomology 34: 649-687 (

Liebherr, J.K. 2008. Taxonomic revision of Hawaiian Bembidion Latreille (Coleoptera: Carabidae, Bembidiini) with a discussion of their reductive and derivative evolutionary specializations. Annals of Carnegie Museum 77: 31-78 (

Liebherr, J.K. 2008. Taxonomic revision of Mecyclothorax Sharp (Coleoptera: Carabidae) of Hawaii Island: abundant genitalic variation in a nascent island radiation. Deutsche Entomologische Zeitshrift 55: 19-78 (

Short, A.E.Z. & J.K. Liebherr. 2007. Systematics and biology of the endemic water scavenger beetles of Hawaii (Coleoptera: Hydrophilidae, Hydrophilini). Systematic Entomology 32: 601-624 (

Liebherr, J.K. 2007[2006]. Taxonomic revision of the Mecyclothorax beetles (Coleoptera: Carabidae, Psydrini) of Molokai, Hawaii and recognition of areas of endemism on Kamakou Volcano. Journal of the New York Entomological Society 114: 179-281 (

Liebherr, J.K. 2006. Hawaiian Blackburnia beetles (Coleoptera, Carabidae, Platynini): patterns of specialization with implications for conservation. Mitteilungen der Deutsche Gesellschaft fur Allgemeine und Angewandte Entomologie 15: 57-62 (pdf).

Liebherr, J.K. 2006. Recognition and description of Blackburnia kavanaughi, new species (Coleoptera: Carabidae) from Kauai, Hawaii. Journal of the New York Entomological Society 114:17-27 (

Liebherr, J.K. & A.E.Z. Short. 2006. Blackburnia riparia, new species (Coleoptera: Carabidae, Platynini): a novel element in the Hawaiian riparian insect fauna. Journal of the New York Entomological Society 114:1-16 ( For data matrix used in this paper and Liebherr [2006], and the 10,108 MEPTs found, see: data matrix.

Liebherr, J.K. & P.D. Krushelnycky. 2006. Unfortunate encounters? Novel interactions of native Mecyclothorax, alien Trechus obtusus(Coleoptera: Carabidae) and Agentine ant (Linepithema humile, Hymenoptera: Formicidae) across a Hawaiian landscape. Journal of Insect Conservation 11: 61-73 (

Liebherr, J.K. 2006[2005]. New species of Mecyclothorax (Coleoptera: Carabidae, Psydrini) from Polipoli, Maui define an area of endemism on Haleakala Volcano, Hawaii. Journal of the New York Entomological Society 113: 97-128 (

Liebherr, J.K. 2005. Vanuatu Platynini (Coleoptera: Carabidae): Miocene diversification on the Melanesian Arc. Invertebrate Systematics 19: 263-295 (

Krushelnycky, P.D., R.G. Gillespie, L.L. Loope & J.K. Liebherr. 2005. Rediscovery and uncertain future of high-elevation Haleakalā carabid beetles (Coleoptera). Pacific Science 59: 399-410 (

Liebherr, J.K. 2004. Patterns of endangerment or pathways to enlightenment? Reconstructing the Fauna Hawaiiensis. Systematics & Biodiversity 2: 175-189 ( 

Polhemus, D.A., C.P. Ewing, R. Kaholoa`a & J.K. Liebherr. 2003. Rediscovery of Blackburnia anomala (Blackburn) (Coleoptera: Carabidae) in East Maui, Hawaii after a 107-year hiatus. Pacific Science 57: 161-165 (

Liebherr, J.K. 2003. Blackburnia lata sp. n. (Coleoptera: Carabidae) from Kauai: morphological transformation in the arboreal microhabitat. Insect Systematics & Evolution 34:41-52 (

Liebherr, J.K. & R. Takumi. 2002. Introduction and distributional expansion of Trechus obtusus Erichson (Coleoptera: Carabidae) in Maui, Hawaii. Pacific Science 56: 365-375 (

Liebherr, J.K. 2001. Blackburnia gastrellariformis, sp. n. (Coleoptera: Carabidae) from Molokai: successful prediction of a new taxon by reconciled tree analysis. Insect Systematics & Evolution 32:133-141 (×00083).

Cryan, J.R., J.K. Liebherr, J.W. Fetzner, Jr. & M.F. Whiting. 2001. Evaluation of relationships within the endemic Hawaiian Platynini (Coleoptera: Carabidae) based on molecular and morphological evidence. Molecular Phylogenetics & Evolution 19: 72-85 (

Liebherr, J.K. & E.C. Zimmerman. 2000. Hawaiian Carabidae (Coleoptera), Part 1: Introduction and Tribe Platynini. Insects of Hawaii 16: v + 494 pp. University of Hawaii Press, Honolulu (

Liebherr, J.K. & E.C. Zimmerman. 1998. Cladistic analysis, phylogeny, and biogeography of Hawaiian Platynini (Coleoptera: Carabidae). Systematic Entomology 23: 137-172 (10.1046/j.1365-3113.1998.00044.x).

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